List of Publications (Google Scholar Citations


[61] K. Ishimoto, C. Moreau and J. Herault

Robust undulatory locomotion via neuromechanical adjustments in a dissipative medium

Journal of the Royal Society Interface, to appear.


[60] S. Unnikrishnan, R. L. Scott, E, Ogundele, M. A. Azad, K. Ishimoto, S. S. Suarez and C. K. Tung

Hybrid motility mechanism of sperm at viscoelastic fluid-solid interface

Scientific Reports, 14 (2024) 21841.

doi: 10.1038/s41598-024-72816-y


[59] K. Yasuda, K. Ishimoto and S. Komura

Statistical formulation of the Onsager-Machlup variational principle

Physical Review E, 110 (2024) 044104.

doi: 10.1103/PhysRevE.110.044104


[58] K. Ogawa and K. Ishimoto

Generalised Taylor dispersion of chiral microswimmers

Philosophical Transactions of the Royal Society A, to appear.


[57] L.-S. Lin, K. Yasuda, K. Ishimoto and S. Komura

Emergence of odd elasticity in a microswimmer using deep reinforcement learning

Physical Review Research, 6 (2024) 033016. 

doi: 10.1103/PhysRevResearch.6.033016


[56] K. Ishimoto (in Japanese)

Continuum mechanics of living matter: A tale of tales in cellular swimming

Butsuri, 79 (2024) 486-494.

doi: 10.11316/butsuri.79.9_486


[55] Y. Hiruta and K. Ishimoto

Reciprocal microswimming in fluctuating and confined environments

Physical Review Research, 6 (2024) 013117.

doi: 10.1103/PhysRevResearch.6.013117


[54] M. P. Dalwadi, C. Moreau, E. A. Gaffney, B. J. Walker and K. Ishimoto

Generalised Jeffery's equations for rapidly spinning particles. Part 2: Helicoidal objects with chirality

Journal of Fluid Mechanics, 979 (2024) A2. 

doi: 10.1017/jfm.2023.924


[53] M. P. Dalwadi, C. Moreau, E. A. Gaffney, K. Ishimoto and B. J. Walker

Generalised Jeffery's equations for rapidly spinning particles. Part 1: Spheroids

Journal of Fluid Mechanics, 979 (2024) A1. 

doi: 10.1017/jfm.2023.923


[52] K. Ishimoto and M. Iima

Preface: Advances in the physics of biofluid locomotion

Journal of the Physical Society of Japan, 92(12) (2023). (pdf)

Featured in JPS Hot Topics “Exploring recent advances in the physics of biofluid locomotion” 


[51] K. Ishimoto, C. Moreau and K. Yasuda

Odd elastohydrodynamics: Non-reciprocal living material in a viscous fluid

PRX Life, 1 (2023) 023002. 

doi: 10.1103/PRXLife.1.023002

Featured in New Scientist “Sperm caught breaking Newton's third law of motion”

Selected as an Invited Talk in APS March Meeting 2024


[50] A. Kobayashi, K. Yasuda, K. Ishimoto, L.-S. Lin, I. Sou, Y. Hosaka and S. Komura

Odd elasticity of a catalytic micromachine

Journal of the Physical Society of Japan,  92 (2023) 074801.

doi: 10.7566/JPSJ.92.074801


[49] K. Ishimoto

Jeffery's orbits and microswimmers in flows: A theoretical review

Journal of the Physical Society of Japan, 92 (2023) 062001.

doi: 10.7566/JPSJ.92.062001

Featured in JPS Hot Topics “Microswimmer hydrodynamics through the lens of Jeffery's orbits”


[48] K. Ishimoto, E. A. Gaffney and D. J. Smith

Squirmer hydrodynamics near a periodic surface topography

Frontiers in Cell and Developmental Biology, 11 (2023) 1123446.

doi: 10.3389/fcell.2023.1123446


[47] B. J. Walker, K. Ishimoto and E. A. Gaffney

Systematic parameterization for minimal models of microswimming

Physical Review Fluids, 8 (2023) 034102.

doi: 10.1103/PhysRevFluids.8.034102


[46] B. J. Walker, K. Ishimoto and E. A. Gaffney

A hydrodynamic slender-body theory for local rotation at zero Reynolds number

Physical Review Fluids, 8 (2023) 034101.

doi: 10.1103/PhysRevFluids.8.034101


[45] L.-S. Lin, K. Yasuda, K. Ishimoto, Y. Hosaka and S. Komura

Onsager's variational principle for nonreciprocal system with odd elasticity

Journal of the Physical Society of Japan, 92 (2023) 033001. 

doi: 10.7566/JPSJ.92.033001


[44] K. Yasuda and K. Ishimoto

The most probable path of an active Brownian particle

Physical Review E, 106 (2022) 064120. 

doi: 10.1103/PhysRevE.106.064120


[43] K. Ishimoto

A multi-scale numerical simulation of quasi-two-dimensional bacterial turbulence using a regularized Stokeslet representation

Springer Proceedings in Mathematics & Statistics, 429 (2023) 215-226. 

doi: 10.1007/978-3-031-35871-5_11 


[42] T. Ayukawa, M. Akiyama, Y. Hozumi, K. Ishimoto, J. Sasaki, H. Senoo, T. Sasaki and M. Yamazaki

Tissue flow regulates planar cell polarity independently of the Frizzled core pathway

Cell Reports, 40 (2022) 111388. 

doi: 10.1016/j.celrep.2022.111388


[41] K. Yasuda, K. Ishimoto, A. Kobayashi, L.-S. Lin, I. Sou, Y. Hosaka and S. Komura

Time-correlation functions for odd Langevin systems

The Journal of Chemical Physics, 157 (2022) 095101. 

doi: 10.1063/5.0095969


[40] B. J. Walker, K. Ishimoto, C. Moreau, E. A. Gaffney and M. P. Dalwadi 

Emergent rheotaxis of shape-changing swimmers in Poiseuille flow

Journal of Fluid Mechanics, 944 (2022) R2. 

doi: 10.1017/jfm.2022.474

Featured in Focus on Fluids ”When do shape changers swim upstream?”


[39] K. Ishimoto, C. Moreau and K. Yasuda

Self-organized swimming with odd elasticity

Physical Review E, 105 (2022) 064603.

doi: 10.1103/PhysRevE.105.064603

Featured in EurekAlert! “Never too odd to learn how to swim”

Featured in Science Japan “Theory of self-organized swimming with odd elasticity discovered”

Featured in Nagare: Journal of Japan Society of Fluid Mechanics, 41 (2022) 411-414.


[38] B. J. Walker, K. Ishimoto,  E. A. Gaffney and C. Moreau

The control of particles in the Stokes limit

Journal of Fluid Mechanics, 942 (2022) A1.

doi: 10.1017/jfm.2022.253


[37] E. A. Gaffney, M. P. Dalwadi, C. Moreau, K. Ishimoto and  B. J. Walker

Canonical orbits for rapidly deforming planar microswimmers in shear flow

Physical Review Fluids, 7 (2022) L022101.

doi: 10.1103/PhysRevFluids.7.L022101


[36] B. J. Walker, K. Ishimoto, E. A. Gaffney, C. Moreau and M. P. Dalwadi 

The effects of rapid yawing on simple swimmer models and planar Jeffery's orbits

Physical Review Fluids, 7 (2022) 023101. 

doi: 10.1103/PhysRevFluids.7.023101


[35] C. Moreau and K. Ishimoto

Driving a microswimmer with wall-induced flow

Micromachines, 12 (2021) 1025. 

doi: 10.3390/mi12091025


[34] C. Moreau, K. Ishimoto, E. A. Gaffney and B. J. Walker

Control and controllability of microswimmers by a shearing flow

Royal Society Open Science, 8 (2021) 211141.

doi: 10.1098/rsos.211141


[33] E. A. Gaffney, K. Ishimoto and B. J. Walker

Modelling motility: the mathematics of spermatozoa

Frontiers in Cell and Developmental Biology, 9 (2021) 710825.

doi: 10.3389/fcell.2021.710825


[32] B. J. Walker, K. Ishimoto and E. A. Gaffney

Efficient simulation of filament elastohydrodynamics in three dimensions

Physical Review Fluids, 5 (2020) 123103.

doi: 10.1103/PhysRevFluids.5.123103

Selected as Editors' Suggestion.


[31] K. Ishimoto, E. A. Gaffney and B. J. Walker

Regularized representation of bacterial hydrodynamics

Physical Review Fluids, 5 (2020) 093101.

doi: 10.1103/PhysRevFluids.5.093101


[30] K. Ishimoto

Jeffery orbits for an object with discrete rotational symmetry

Physics of Fluids, 32 (2020) 081904.

doi: 10.1063/5.0015056


[29] B. J. Walker, M. P. Curtis, K. Ishimoto and E. A. Gaffney

A regularised slender-body theory of non-uniform filaments

Journal of Fluid Mechanics, 899 (2020) A3.

doi: 10.1017/jfm.2020.434


[28] B. J. Walker, S. Phuyal, K. Ishimoto, C.-K. Tung and E. A. Gaffney

Computer-assisted beat-pattern analysis and the flagellar waveforms of bovine spermatozoa

Royal Society Open Science, 7 (2020) 200769. 

doi: 10.1098/rsos.200769


[27] K. Ishimoto

Helicoidal particles and swimmers in a flow at low Reynolds number

Journal of Fluid Mechanics, 892 (2020) A11.

doi: 10.1017/jfm.2020.142


[26] B. J. Walker, K. Ishimoto, H. Gadêlha and E. A. Gaffney

Filament mechanics in a half-space via regularised Stokeslet segments

Journal of Fluid Mechanics, 879 (2019) 808-833.

doi: 10.1017/jfm.2019.723


[25] K. Ishimoto

Bacterial spinning top

Journal of Fluid Mechanics, 880 (2019) 620-652.

doi: 10.1017/jfm.2019.714


[24] B. J. Walker, K. Ishimoto and E. A. Gaffney

Pairwise hydrodynamic interactions of synchronized spermatozoa

Physical Review Fluids, 4 (2019) 093101.

doi: 10.1103/PhysRevFluids.4.093101


[23] K. Ishimoto and E. Lauga

The N-flagella problem: Elastohydrodynamic motility transition of multi-flagellated bacteria

Proceedings of the Royal Society A, 475 (2019) 20180690.

doi: 10.1098/rspa.2018.0690

Featured in Nagare: Journal of Japan Society of Fluid Mechanics, 38 (2019) 407-410.


[22] B. J. Walker, K. Ishimoto and  R. J. Wheeler

Automated identification of flagella from videomicroscopy via the medial axis transform

Scientific Reports, 9 (2019) 5015.

doi: 10.1038/S41598-019-41459-9


[21] B. J. Walker, K. Ishimoto, R. J. Wheeler and E. A. Gaffney

Response of monoflagellate pullers to a shearing flow: A simulation study of microswimmer guidance

Physical Review E, 98 (2018) 063111.

doi: 10.1103/PhysRevE.98.063111


[20] B. J. Walker, R. J. Wheeler, K. Ishimoto and E. A. Gaffney

Boundary behaviours of Leishmania mexicana: a hydrodynamic simulation study

Journal of Theoretical Biology, 462 (2019) 311-320.

doi: 10.1016/j.jtbi.2018.11.016


[19] K. Ishimoto and E. A. Gaffney

Hydrodynamic clustering of human sperm in viscoelastic fluids

Scientific Reports, 8 (2018) 15600.

doi: 10.1038/s41598-018-33584-8


[18] K. Ishimoto and E. A. Gaffney

An elastohydrodynamical simulation study of filament and spermatozoan swimming driven by internal couples

IMA Journal of Applied Mathematics, 83 (2018) 655-679.

doi: 10.1093/imamat/hxy025


[17] K. Ishimoto, H. Gadêlha, E. A. Gaffney, D. J. Smith and J. Kirkman-Brown

Human sperm swimming in a high viscous mucus analogue

Journal of Theoretical Biology, 446 (2018) 1-10.

doi: 10.1016/j.jtbi.2018.02.013


[16] K. Ishimoto

Guidance of microswimmers by wall and flow: Thigmotaxis and rheotaxis of unsteady squirmers in two and three dimensions

Physical Review E, 96 (2017) 043101.

doi: 10.1103/PhysRevE.96.043103


[15] K. Ishimoto and E. A. Gaffney

Boundary element methods for particles and microswimmers in a linear viscoelastic fluid

Journal of Fluid Mechanics, 831 (2017) 228-251.

doi: 10.1017/jfm.2017.636


[14] K. Ishimoto and D. G. Crowdy

Dynamics of a treadmilling microswimmer near a no-slip wall in simple shear

Journal of Fluid Mechanics, 821 (2017) 647-667.

doi: 10.1017/jfm.2017.220


[13] K. Ishimoto, H. Gadêlha, E. A. Gaffney, D. J. Smith and J. Kirkman-Brown

Coarse-graining the fluid flow around a human sperm

Physical Review Letters, 118 (2017) 124501.

doi: 10.1103/PhysRevLett.118.124501

Featured in BBC News “Sperm swimming technique 'all down to simple maths’”.

Featured in Daily Mail Online “The formula for sperm's success: 'Relatively simple' calculation could treat male infertility”

Featured in The Japan Times “Kyoto University professor does the math and finds that sperm swim to a beat”


[12] K. Ishimoto, M. Ikawa and M. Okabe

The mechanics clarifying counterclockwise rotation of most IVF eggs in mice

Scientific Reports, 7 (2017) 43456.

doi: 10.1038/srep43456


[11] K. Ishimoto and E. A. Gaffney

Mechanical tuning of mammalian sperm behaviour by hyperactivation, rheology and substrate adhesion: a numerical exploration

Journal of the Royal Society Interface, 13 (2016) 20160633.

doi: 10.1098/rsif.2016.0633


[10] K. Ishimoto

Hydrodynamic evolution of sperm swimming: Optimal flagella by a genetic algorithm

Journal of Theoretical Biology, 399 (2016) 166-174.

doi: 10.1016/j.jtbi.2016.03.041


[9] K. Ishimoto, J. Cosson and E. A. Gaffney

A simulation study of sperm motility hydrodynamics near fish eggs and spheres

Journal of Theoretical Biology, 389 (2016) 187-197.

doi: 10.1016/j.jtbi.2015.10.013


[8] K. Ishimoto and E. A. Gaffney

Modelling spermatozoan swimming: its capabilities and limitations for contributing to the understanding of sperm guidance

in “Flagellar Mechanics and Sperm Guidance" (Ed. Jacky Cosson, Bentham Science Publishers)  (2015) 315-348.

doi: 10.2174/97816810812811150101


[7] K. Ishimoto and E. A. Gaffney

Fluid flow and sperm guidance: a simulation study of hydrodynamic sperm rheotaxis

Journal of the Royal Society Interface, 12 (2015) 20150172.

doi: 10.1098/rsif.2015.0172


[6] K. Ishimoto and E. A. Gaffney

A study of spermatozoan swimming stability near a surface

Journal of Theoretical Biology, 360 (2014) 187-199.

doi: 10.1016/j.jtbi.2014.06.034


[5] K. Ishimoto and E. A. Gaffney

Swimming efficiency of spherical squirmers: Beyond the Lighthill theory

Physical Review E, 90 (2014) 012704.

doi: 10.1103/PhysRevE.90.012704


[4] K. Ishimoto and M. Yamada (in Japanese)

A coordinate-based proof of the scallop theorem (Special Review by RONBUN-Prize Award)

Nagare: Journal of Japan Society of Fluid Mechanics, 33 (2014) 237-240.


[3] K. Ishimoto and E. A. Gaffney

Squirmer dynamics near a boundary

Physical Review E, 88 (2013) 062702.

doi: 10.1103/PhysRevE.88.062702


[2] K. Ishimoto

A spherical squirming swimmer in unsteady Stokes flow

Journal of Fluid Mechanics, 723 (2013) 163-189.

doi: 10.1017/jfm.2013.131


[1] K. Ishimoto and M. Yamada

A coordinate-based proof of the scallop theorem

SIAM Journal on Applied Mathematics, 72 (2012) 1686-1694.

doi: 10.1137/110853297


Books


[2] K. Ishimoto (Japanese translation)

Fluid Mechanics: A Very Short Introduction” by Eric Lauga (Iwanami Shoten, Publishers, 2023)

ISBN : 978-4000297233


[1] K. Ishimoto (Book in Japanese)

Microbial Fluid Mechanics: Exploring Motion, Shape, and Flow of Life (Saiensu-Sha, 2022)

ISBN : 978-4-7819-1559-3


MISC


[1] K. Ishimoto

Problems in microswimmer hydrodynamics

RIMS Kokyuroku, 2266 (2023) 131-140.


Media Interview


[4] “Sperm caught breaking Newton's third law of motion”

New Scientist, published online on 20 October 2023 (website)

Published in print, Issue 3462, p14 (28 October 2023) (website)

Selected as New Scientist's most popular story of the year (20 December 2023)


[3] “How do Sperm Swim?”

Fluid Mechanics Blog, Journal of Fluid Mechanics, published on 9 November 2020

 (website) (YouTube)


[2] “Underwater Snakes, Gusty Flying, and Microswimmers”

Video Series of FYFD/Journal of Fluid Mechanics Collaboration, published online on 7 December 2018 (YouTube)


[1] “Kyoto University professor does the math and finds that sperm swim to a beat”

The Japan Times, published online on 24 March 2017 (p3, Newspaper on 25 March) (website)


Copyright (c) Kenta Ishimoto, All Rights Reserved.