論文リスト (Google Scholar Citations


[61] K. Ishimoto, C. Moreau and J. Herault

Robust undulatory locomotion via neuromechanical adjustments in a dissipative medium

Journal of the Royal Society Interface, to appear.


[60] S. Unnikrishnan, R. L. Scott, E, Ogundele, M. A. Azad, K. Ishimoto, S. S. Suarez and C. K. Tung

Hybrid motility mechanism of sperm at viscoelastic fluid-solid interface

Scientific Reports, 14 (2024) 21841.

doi: 10.1038/s41598-024-72816-y


[59] K. Yasuda, K. Ishimoto and S. Komura

Statistical formulation of the Onsager-Machlup variational principle

Physical Review E, 110 (2024) 044104.

doi: 10.1103/PhysRevE.110.044104


[58] K. Ogawa and K. Ishimoto

Generalised Taylor dispersion of chiral microswimmers

Philosophical Transactions of the Royal Society A, to appear.


[57] L.-S. Lin, K. Yasuda, K. Ishimoto and S. Komura

Emergence of odd elasticity in a microswimmer using deep reinforcement learning

Physical Review Research, 6 (2024) 033016. 

doi: 10.1103/PhysRevResearch.6.033016


[56] 石本健太

細胞遊泳から始める「生き物」の力学

日本物理学会誌, 79 (2024) 486-494.

doi: 10.11316/butsuri.79.9_486


[55] Y. Hiruta and K. Ishimoto

Reciprocal microswimming in fluctuating and confined environments

Physical Review Research, 6 (2024) 013117.

doi: 10.1103/PhysRevResearch.6.013117


[54] M. P. Dalwadi, C. Moreau, E. A. Gaffney, B. J. Walker and K. Ishimoto

Generalised Jeffery's equations for rapidly spinning particles. Part 2: Helicoidal objects with chirality

Journal of Fluid Mechanics, 979 (2024) A2. 

doi: 10.1017/jfm.2023.924


[53] M. P. Dalwadi, C. Moreau, E. A. Gaffney, K. Ishimoto and B. J. Walker

Generalised Jeffery's equations for rapidly spinning particles. Part 1: Spheroids

Journal of Fluid Mechanics, 979 (2024) A1. 

doi: 10.1017/jfm.2023.923


[52] K. Ishimoto and M. Iima

Preface: Advances in the physics of biofluid locomotion

Journal of the Physical Society of Japan, 92(12) (2023). (pdf)

研究内容がJPS Hot Topicsで紹介されました:“Exploring recent advances in the physics of biofluid locomotion” 


[51] K. Ishimoto, C. Moreau and K. Yasuda

Odd elastohydrodynamics: Non-reciprocal living material in a viscous fluid

PRX Life, 1 (2023) 023002. 

doi: 10.1103/PRXLife.1.023002

論文がNew Scientistで紹介されました:“Sperm caught breaking Newton's third law of motion”

論文がAPS March Meeting 2024招待講演に選ばれました


[50] A. Kobayashi, K. Yasuda, K. Ishimoto, L.-S. Lin, I. Sou, Y. Hosaka and S. Komura

Odd elasticity of a catalytic micromachine

Journal of the Physical Society of Japan,  92 (2023) 074801.

doi: 10.7566/JPSJ.92.074801


[49] K. Ishimoto

Jeffery's orbits and microswimmers in flows: A theoretical review

Journal of the Physical Society of Japan, 92 (2023) 062001.

doi: 10.7566/JPSJ.92.062001

論文が日本物理学会誌 JPSJの最近の注目論文から」で紹介されました(日本物理学会誌, 78 (2023) 717

論文がJPS Hot Topics紹介されました:“Microswimmer hydrodynamics through the lens of Jeffery's orbits”


[48] K. Ishimoto, E. A. Gaffney and D. J. Smith

Squirmer hydrodynamics near a periodic surface topography

Frontiers in Cell and Developmental Biology, 11 (2023) 1123446.

doi: 10.3389/fcell.2023.1123446


[47] B. J. Walker, K. Ishimoto and E. A. Gaffney

Systematic parameterization for minimal models of microswimming

Physical Review Fluids, 8 (2023) 034102.

doi: 10.1103/PhysRevFluids.8.034102


[46] B. J. Walker, K. Ishimoto and E. A. Gaffney

A hydrodynamic slender-body theory for local rotation at zero Reynolds number

Physical Review Fluids, 8 (2023) 034101.

doi: 10.1103/PhysRevFluids.8.034101


[45] L.-S. Lin, K. Yasuda, K. Ishimoto, Y. Hosaka and S. Komura

Onsager's variational principle for nonreciprocal system with odd elasticity

Journal of the Physical Society of Japan, 92 (2023) 033001. 

doi: 10.7566/JPSJ.92.033001


[44] K. Yasuda and K. Ishimoto

The most probable path of an active Brownian particle

Physical Review E, 106 (2022) 064120. 

doi: 10.1103/PhysRevE.106.064120


[43] K. Ishimoto

A multi-scale numerical simulation of quasi-two-dimensional bacterial turbulence using a regularized Stokeslet representation

Springer Proceedings in Mathematics & Statistics, 429 (2023) 215-226. 

doi: 10.1007/978-3-031-35871-5_11 


[42] T. Ayukawa, M. Akiyama, Y. Hozumi, K. Ishimoto, J. Sasaki, H. Senoo, T. Sasaki and M. Yamazaki

Tissue flow regulates planar cell polarity independently of the Frizzled core pathway

Cell Reports, 40 (2022) 111388. 

doi: 10.1016/j.celrep.2022.111388


[41] K. Yasuda, K. Ishimoto, A. Kobayashi, L.-S. Lin, I. Sou, Y. Hosaka and S. Komura

Time-correlation functions for odd Langevin systems

The Journal of Chemical Physics, 157 (2022) 095101. 

doi: 10.1063/5.0095969


[40] B. J. Walker, K. Ishimoto, C. Moreau, E. A. Gaffney and M. P. Dalwadi 

Emergent rheotaxis of shape-changing swimmers in Poiseuille flow

Journal of Fluid Mechanics, 944 (2022) R2. 

doi: 10.1017/jfm.2022.474

論文がFocus on Fluids紹介されました:When do shape changers swim upstream?


[39] K. Ishimoto, C. Moreau and K. Yasuda

Self-organized swimming with odd elasticity

Physical Review E, 105 (2022) 064603.

doi: 10.1103/PhysRevE.105.064603

論文が京都大学ウェブページで紹介されました:「自ら流体中を泳ぐ「奇弾性体」の発見」

論文がEurekAlert!で紹介されました:「Never too odd to learn how to swim

研究内容が日本流体力学会年会注目研究に選ばれました:「奇弾性体の微小遊泳理論」


[38] B. J. Walker, K. Ishimoto,  E. A. Gaffney and C. Moreau

The control of particles in the Stokes limit

Journal of Fluid Mechanics, 942 (2022) A1.

doi: 10.1017/jfm.2022.253


[37] E. A. Gaffney, M. P. Dalwadi, C. Moreau, K. Ishimoto and  B. J. Walker

Canonical orbits for rapidly deforming planar microswimmers in shear flow

Physical Review Fluids, 7 (2022) L022101.

doi: 10.1103/PhysRevFluids.7.L022101


[36] B. J. Walker, K. Ishimoto, E. A. Gaffney, C. Moreau and M. P. Dalwadi 

The effects of rapid yawing on simple swimmer models and planar Jeffery's orbits

Physical Review Fluids, 7 (2022) 023101. 

doi: 10.1103/PhysRevFluids.7.023101


[35] C. Moreau and K. Ishimoto

Driving a microswimmer with wall-induced flow

Micromachines, 12 (2021) 1025. 

doi: 10.3390/mi12091025


[34] C. Moreau, K. Ishimoto, E. A. Gaffney and B. J. Walker

Control and controllability of microswimmers by a shearing flow

Royal Society Open Science, 8 (2021) 211141.

doi: 10.1098/rsos.211141


[33] E. A. Gaffney, K. Ishimoto and B. J. Walker

Modelling motility: the mathematics of spermatozoa

Frontiers in Cell and Developmental Biology, 9 (2021) 710825.

doi: 10.3389/fcell.2021.710825


[32] B. J. Walker, K. Ishimoto and E. A. Gaffney

Efficient simulation of filament elastohydrodynamics in three dimensions

Physical Review Fluids, 5 (2020) 123103.

doi: 10.1103/PhysRevFluids.5.123103

論文がEditors' Suggestionに選ばれました


[31] K. Ishimoto, E. A. Gaffney and B. J. Walker

Regularized representation of bacterial hydrodynamics

Physical Review Fluids, 5 (2020) 093101.

doi: 10.1103/PhysRevFluids.5.093101


[30] K. Ishimoto

Jeffery orbits for an object with discrete rotational symmetry

Physics of Fluids, 32 (2020) 081904.

doi: 10.1063/5.0015056


[29] B. J. Walker, M. P. Curtis, K. Ishimoto and E. A. Gaffney

A regularised slender-body theory of non-uniform filaments

Journal of Fluid Mechanics, 899 (2020) A3.

doi: 10.1017/jfm.2020.434


[28] B. J. Walker, S. Phuyal, K. Ishimoto, C.-K. Tung and E. A. Gaffney

Computer-assisted beat-pattern analysis and the flagellar waveforms of bovine spermatozoa

Royal Society Open Science, 7 (2020) 200769. 

doi: 10.1098/rsos.200769


[27] K. Ishimoto

Helicoidal particles and swimmers in a flow at low Reynolds number

Journal of Fluid Mechanics, 892 (2020) A11.

doi: 10.1017/jfm.2020.142


[26] B. J. Walker, K. Ishimoto, H. Gadêlha and E. A. Gaffney

Filament mechanics in a half-space via regularised Stokeslet segments

Journal of Fluid Mechanics, 879 (2019) 808-833.

doi: 10.1017/jfm.2019.723


[25] K. Ishimoto

Bacterial spinning top

Journal of Fluid Mechanics, 880 (2019) 620-652.

doi: 10.1017/jfm.2019.714

論文が京都大学ウェブページで紹介されました:「微生物の運動に潜む柔らかな力学メカニズムを解明- バクテリア「独楽」は如何にして廻るか-


[24] B. J. Walker, K. Ishimoto and E. A. Gaffney

Pairwise hydrodynamic interactions of synchronized spermatozoa

Physical Review Fluids, 4 (2019) 093101.

doi: 10.1103/PhysRevFluids.4.093101


[23] K. Ishimoto and E. Lauga

The N-flagella problem: Elastohydrodynamic motility transition of multi-flagellated bacteria

Proceedings of the Royal Society A, 475 (2019) 20180690.

doi: 10.1098/rspa.2018.0690

研究内容が日本流体力学会年会注目研究に選ばれました:「N鞭毛問題:バクテリア遊泳の安定性」

研究内容の図が日本流体力学会誌「ながれ」38(2019)6表紙選ばれました


[22] B. J. Walker, K. Ishimoto and  R. J. Wheeler

Automated identification of flagella from videomicroscopy via the medial axis transform

Scientific Reports, 9 (2019) 5015.

doi: 10.1038/S41598-019-41459-9


[21] B. J. Walker, K. Ishimoto, R. J. Wheeler and E. A. Gaffney

Response of monoflagellate pullers to a shearing flow: A simulation study of microswimmer guidance

Physical Review E, 98 (2018) 063111.

doi: 10.1103/PhysRevE.98.063111


[20] B. J. Walker, R. J. Wheeler, K. Ishimoto and E. A. Gaffney

Boundary behaviours of Leishmania mexicana: a hydrodynamic simulation study

Journal of Theoretical Biology, 462 (2019) 311-320.

doi: 10.1016/j.jtbi.2018.11.016


[19] K. Ishimoto and E. A. Gaffney

Hydrodynamic clustering of human sperm in viscoelastic fluids

Scientific Reports, 8 (2018) 15600.

doi: 10.1038/s41598-018-33584-8


[18] K. Ishimoto and E. A. Gaffney

An elastohydrodynamical simulation study of filament and spermatozoan swimming driven by internal couples

IMA Journal of Applied Mathematics, 83 (2018) 655-679.

doi: 10.1093/imamat/hxy025


[17] K. Ishimoto, H. Gadêlha, E. A. Gaffney, D. J. Smith and J. Kirkman-Brown

Human sperm swimming in a high viscous mucus analogue

Journal of Theoretical Biology, 446 (2018) 1-10.

doi: 10.1016/j.jtbi.2018.02.013


[16] K. Ishimoto

Guidance of microswimmers by wall and flow: Thigmotaxis and rheotaxis of unsteady squirmers in two and three dimensions

Physical Review E, 96 (2017) 043101.

doi: 10.1103/PhysRevE.96.043103


[15] K. Ishimoto and E. A. Gaffney

Boundary element methods for particles and microswimmers in a linear viscoelastic fluid

Journal of Fluid Mechanics, 831 (2017) 228-251.

doi: 10.1017/jfm.2017.636


[14] K. Ishimoto and D. G. Crowdy

Dynamics of a treadmilling microswimmer near a no-slip wall in simple shear

Journal of Fluid Mechanics, 821 (2017) 647-667.

doi: 10.1017/jfm.2017.220


[13] K. Ishimoto, H. Gadêlha, E. A. Gaffney, D. J. Smith and J. Kirkman-Brown

Coarse-graining the fluid flow around a human sperm

Physical Review Letters, 118 (2017) 124501.

doi: 10.1103/PhysRevLett.118.124501

論文が京都大学ウェブページで紹介されました:「数理の目で見る精子の旅」

論文がBBC News で紹介されました:“Sperm swimming technique 'all down to simple maths’”.

インタビュー記事が The Japan Times に掲載されました:“Kyoto University professor does the math and finds that sperm swim to a beat”.


[12] K. Ishimoto, M. Ikawa and M. Okabe

The mechanics clarifying counterclockwise rotation of most IVF eggs in mice

Scientific Reports, 7 (2017) 43456.

doi: 10.1038/srep43456


[11] K. Ishimoto and E. A. Gaffney

Mechanical tuning of mammalian sperm behaviour by hyperactivation, rheology and substrate adhesion: a numerical exploration

Journal of the Royal Society Interface, 13 (2016) 20160633.

doi: 10.1098/rsif.2016.0633


[10] K. Ishimoto

Hydrodynamic evolution of sperm swimming: Optimal flagella by a genetic algorithm

Journal of Theoretical Biology, 399 (2016) 166-174.

doi: 10.1016/j.jtbi.2016.03.041


[9] K. Ishimoto, J. Cosson and E. A. Gaffney

A simulation study of sperm motility hydrodynamics near fish eggs and spheres

Journal of Theoretical Biology, 389 (2016) 187-197.

doi: 10.1016/j.jtbi.2015.10.013


[8] K. Ishimoto and E. A. Gaffney

Modelling spermatozoan swimming: its capabilities and limitations for contributing to the understanding of sperm guidance

in “Flagellar Mechanics and Sperm Guidance" (Ed. Jacky Cosson, Bentham Science Publishers)  (2015) 315-348.

doi: 10.2174/97816810812811150101


[7] K. Ishimoto and E. A. Gaffney

Fluid flow and sperm guidance: a simulation study of hydrodynamic sperm rheotaxis

Journal of the Royal Society Interface, 12 (2015) 20150172.

doi: 10.1098/rsif.2015.0172


[6] K. Ishimoto and E. A. Gaffney

A study of spermatozoan swimming stability near a surface

Journal of Theoretical Biology, 360 (2014) 187-199.

doi: 10.1016/j.jtbi.2014.06.034


[5] K. Ishimoto and E. A. Gaffney

Swimming efficiency of spherical squirmers: Beyond the Lighthill theory

Physical Review E, 90 (2014) 012704.

doi: 10.1103/PhysRevE.90.012704


[4] 石本健太, 山田道夫 

座標に基づいた帆立貝定理の証明

日本流体力学会誌「ながれ」, 33 (2014) 237-240. (pdf)


[3] K. Ishimoto and E. A. Gaffney

Squirmer dynamics near a boundary

Physical Review E, 88 (2013) 062702.

doi: 10.1103/PhysRevE.88.062702


[2] K. Ishimoto

A spherical squirming swimmer in unsteady Stokes flow

Journal of Fluid Mechanics, 723 (2013) 163-189.

doi: 10.1017/jfm.2013.131


[1] K. Ishimoto and M. Yamada

A coordinate-based proof of the scallop theorem

SIAM Journal on Applied Mathematics, 72 (2012) 1686-1694.

doi: 10.1137/110853297


会議録・講究録


[8] K. Ishimoto

Problems in microswimmer hydrodynamics

数理解析研究所講究録、2266 (2023) 131-140. 


[7] 石本健太, Clément Moreau, 安田健人

奇弾性体の微小遊泳理論

日本流体力学会誌「ながれ」, 41 (2022) 411-414. (pdf)


[6] 石本健太, Eric Lauga

N鞭毛問題:バクテリア遊泳の安定性

日本流体力学会誌「ながれ」, 38 (2019) 407-410. (pdf)

研究内容の図が表紙に選ばれました


[5] 石本健太

微生物遊泳における計測問題

数理解析研究所講究録, 1940 (2015) 1-15. (pdf)

hdl: 2433/223797


[4] 石本健太

精子の遊泳と境界の相互作用

数理解析研究所講究録, 1900 (2014) 145-149. (pdf)

hdl: 2433/195916


[3] 石本健太, 小泉吉輝, 鈴木理

肝臓におけるアルコール代謝の数理モデリング:肝障害の発生メカニズムとその予防策

数理解析研究所講究録, 1863 (2013) 29-37. (pdf)

hdl: 2433/195329


[2] 石本健太

球形微生物の運動における慣性の影響

数理解析研究所講究録, 1808 (2012) 56-72. (pdf)

hdl: 2433/194447


[1] 石本健太、山田道夫

微生物の運動と流体力学:帆立貝定理とその破れ

数理解析研究所講究録, 1796 (2012) 15-21. (pdf)

hdl: 2433/172918



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